Bachelor of Arts from the Indiana University, 1959; Ph.D. from the Indiana University, 1963; semi finalist in NASA group 4 selection; 1964 - 1970 Assistant Professor at the Department of Zoology, University of California in Berkeley; 1996 Professor at the Department of Genetics (now Molecular Genetics), Ohio State University; 1997-2007 Professor, Department of Ecology and Evolutionary Biology, University of Arizona; 1997-2003 Chair, Graduate Interdisciplinary Program in Genetics; since 2007 Professor Emeritus. Elected President of the American Genetics Association, 1992-1993. Elected Fellow of the American Association for the Advancement of Science in 1997. Author of more than 75 scientific publications, still active in research.