Bachelor of Science in Biochemistry, University of Surrey, 1979 - 1982; Master of Science in Neurochemistry, University of London, 1982 - 1983; Ph.D. in Neuropharmacology, University of Bath, 1983 - 1986; 1986 - 1987 Research Officer, School of Biological Sciences, University of Surrey, Guild-ford, Surrey, UK; 1987-1990 Postdoctoral Research Fellow, School of Biological Sciences, University of Surrey, Guildford, Surrey, UK; 1990 – 1993 Postdoctoral Research Fellow, Department of Pharmacology, University of Rochester, Rochester, NY, USA; 1994 – 1997 Senior Research Toxicologist, Institute of Occupational Medicine, Edinburgh; 1997 – 2004 Principal Scientist, Exposure Modelling Section, Health Sciences Group, Health & Safety Laboratory, Sheffield; 2004 – 2006 Head, Computational Toxicology Section, Health Sciences Group, Health & Safety Laboratory, Sheffield; since 2006 Head, Computational Toxicology Team, Mathematical Sciences Unit, Health Improvement Group, Health & Safety Laboratory, Buxton; candidate for the Juno project, but did not reach the final selection